Picture with logotype for scandinavian entomology


Entomologica Scandinavica Supplement 51—60

now changed to Insect Systematics and Evolution Supplements


ESS 51.
ENGHOFF, H. (Ed.) Many-legged animals — A collection of papers on Myriapoda and Onychophora. Proceeding of the 10th Intern. Congr. in 1996. 1997. 329 pp. SEK 240.

Enghoff, H.: Introduction: Myriapods and myriapodology.
Dohle, W.: Are the insects more closely related to the crustaceans than to the myriapods?
Rosenberg, J., Krüger, E. & Peters, W.: Intense receptor-mediated endocytosis in nephrocytes of Myriapoda.
Shultz, J. W. & Regier, J. C.: Progress towards a molecular phylogeny of the centipede orders (Chilopoda).
Schileyko, A. A. & Pavlinov, I. J.: A cladistic analysis of the order Scolopendromorpha (Chilopoda).
Prunescu, C.-C.: The anatomy and evolution of the genital system in Scolopendromorpha (Chilopoda).
Hilken, G.: Tracheal systems in Chilopoda: a comparison under phylogenetic aspects.
Berto, D., Fusco, G. & Minelli, A.: Segmental units and shape control in Chilopoda.
Greven, H., Rosenberg, J. & Latka, I.: Cytochemical notes on the specialized cuticle of the coxal organs in +Lithobius forficatus: Demonstration of lecitins and chitin (Chilopoda, Lithobiomorpha: Lithobiidae).
Pereira, L. A., Foddai, D. & Minelli, A.: Zoogeographical aspects of Neotropical Geophilomorpha (Chilopoda).
Stoev, P.: A check-list of the centipedes of the Balkan peninsula with some taxonomic notes and a complete bibliography (Chilopoda).
Fründ, H.-C., Balkenhol, B. & Ruszkowski, B.: Chilopoda in forest habitat-islands in north-west Westphalia, Germany.
Morais, J. W. de, Adis, J., Berti-Filho, E., Pereira, L. A., Minelli, A. & Barbieri, F.: On abundance, phenology and natural history of Geophilomorpha from a mixedwater inundation forest in Central Amazonia (Chilopoda).
Zapparoli, M.: Centipedes of a wasteland urban area in Rome, Italy (Chilopoda).
Zerm, M.: Distribution and phenology of Lamyctes fulvicornis and other lithobiomorph centipedes in the floodplain of the Lower Oder Valley, Germany (Chilopoda: Henicopidae, Lithobiidae).
Hoess, R., Scholl, A. & Lörtscher, M.: The Glomeris-taxa hexasticha and intermedia: species or subspecies? — allozyme data (Diplopoda, Glomerida: Glomeridae).
Mauriès, J.-P.: Is the family Atopogestidae based on a case of teratology or a periodomorphic stage? (Diplopoda, Spirostreptida, Odontopygoidea).
Sørensen, L.: A new species of the previously monotypic genus Allocotoproctus from the Uluguru Mountains, Tanzania (Diplopoda, Polydesmida: Oxydesmidae).
Bano, K. & Murthy, J. B.: Gonopods of some South Indian paradoxosomatid millipedes (Diplopoda, Polydesmida).
Curcic, B. P. & Makarov, S. E.: Postembryonic development of Bulgarosoma lazarevensis — a cave dwelling milliped from Yugoslavia (Diplopoda, Chordeumatida: Anthroleucosomatidae).
Nguyen Duy-Jacquemin, M.: Fine structure and possible functions of antennal sensilla in Polyxenus lagurus (Diplopoda, Penicillata: Polyxenidae).
Kania, G. & Rzeski, W.: In vitro phagocytic activity of hemocytes of Ommatoiulus sabulosus: preliminary observations (Diplopoda, Julida: Julidae).
Xylander, W. E. R. & Bogusch, O.. Granular hemocytes as the main location of prophenoloxidase in the millipede Rhapidostreptus virgator (Diplopoda, Spirostreptida: Spirostreptidae).
Curcic, B. P. & Makarov, S. E.: Diversification and biogeographic features in millipedes in Serbia, Yugoslavia (Diplopoda).
Golovatch, S. I.: On the main traits of milliped distribution and faunogenesis in Eurasia (Diplopoda).
Hamer, L. A.: A preliminary assessment of the southern African milliped fauna: diversity and conservation (Diplopoda).
Korsos, Z.: The milliped fauna of the Dr‡va region, southern Hungary (Diplopoda).
Tajovsky, K.: Distribution of millipedes along an altitudinal gradient in three mountain regions in the Czech and Slovak Republics (Diplopoda).
Wytwer, J. Millipede communities of inundated ash-alder forests in Puszcza Bialowieska, Poland (Diplopoda).
Bailey, P. T.: Decline of an invading millipede, Ommatoiulus moreleti in South Australia: the need for a better understanding of the mechanism (Diplopoda, Julida: Julidae).
Bourdanné, D. K.: Influence de l´anthropisation sur le peuplement en Diplopodes de la région d´Abidjan, Côte d´Ivoire (Diplopoda).
David, J.-F. & Vannier, G.: Cold-hardiness of European millipedes (Diplopoda).
Ishii, K.: Comparative biological study of the penicillate diplopods in Japan (Diplopoda, Penicillata).
Kime, R. D.: Year-round pitfall trapping of millipedes in mainly open grassland in Belgium (Diplopoda).
Mahsberg, D.: Pelmatojulus tigrinus, a key detritivore of a tropical gallery forest (Diplopoda, Spirobolida: Pachybolidae).
Krell, F.-T., Schmitt, T. & Linsenmair, K. E.: Diplopod defensive secretions as attractants for necrophagous scarab beetles (Diplopoda; Insecta, Coleoptera: Scarabaeidae).
Mwabvu, T.: Millipedes in small-scale farming systems in Zimbabwe: abundance and diversity (Diplopoda, Spirostreptida).
Serra, A., Miquel, C., Mateos, E. & Vicente, C.: Study of a soil Julidae community in Mediterranean forest (Diplopoda, Julidae).
Adis, J., Caoduro, G., Messner, B. & Enghoff, H.: On the semiaquatic behaviour of a new troglobitic millipede from northern Italy (Diplopoda, Polydesmida: Polydesmidae).
Adis, J., Scheller, U., Morais, J. W. de, Rochus, C. & Rodrigues, J. M. G.: Symphyla from Amazonian non-flooded upland forests and their adaptations to inundation forests.
Brockmann, C., Mesibov, R. & Ruhberg, H.: Observations on Ooperipatellus decoratus, an oviparous onychophoran from Tasmania (Onychophora: Peripatopsidae).


ESS 52.
HEDQVIST, K.-J. Bark beetles enemies in Sweden. II. Braconidae (Hymenoptera). 1998. 87 pp. SEK 150.

The paper is a result of an inventory work on braconid enemies to Swedish bark beetle species. It was carried out in 1979 to 1989 and my own collections for this purpose covers most of the Swedish landscapes (see map). As a comparison to Swedish conditions an account is given to European investigations and conditions in this field. Also, information on braconids and their parasitoids is extracted from the literature and a table of valid names and synonyms is presented. Braconid morphology is briefly treated to enlighten terms used in the keys and descriptions.


A chapter is devoted to braconid biology, behaviour, mating and methods for finding hosts for oviposition. Species known to occur in Sweden are listed together with their hosts. Methods for collecting, rearing and preparation are shortly described and illustrated. Under the heading ”Taxonomy” all braconid enemies to bark beetles known to occur in Sweden are treated: synonymies, morphological characteristics, hosts and known distribution are given. Keys to all subfamilies, genera and species are provided. The information is summarized in a table showing all Swedish bark beetle species with their braconid parasitoids.


A chapter is devoted to the braconid enemies. Also, the importance of Braconidae to forestry is discussed showing their role in controlling bark beetles. A summary in Swedish is given.


ESS 53.
JOHANSON, K.-A. Phylogenetic and biogeographic analysis of the family Helicopsychidae (Insecta: Trichoptera). 1998. 172 pp. SEK 240

ESS 54.
LUNDQVIST, L. Taxonomic revision of the genus Dinogamasus Kramer (Acari Mesostigmata: Laelapidae). 1999. 109 pp. SEK 160.

The genus Dinogamasus (Kramer, 1898) is taxonomically revised and phylogenetically analysed. There are altogether 37 valid names assigned to the genus prior to this investigation.


Eight species are described as new to science: Dinogamasus albulus n.sp., D. assimiensis n.sp., D. kordofaniensis n.sp., D. levequae n.sp., D. minor n.sp., D. occidentalis n.sp., D. tonkinensis n.sp., and D. tortivus n.sp. All known type material have been investigated, except for the 4 Berlese species [Greenia hirtissima 1910 (?= Dinogamasus inflatus LeVeque 1930?), G. jacobsoni 1910 (= D. longipes LeVeque, 1931), Dolaea affinis 1918 (= Dinogamasus cockerelli LeVeque, 1930; D. northolmensis Loots, 1980) and D. villosior 1918]. Greenia sjoestedti Trägårdh, 1904 and Dolaea schoutedeni Oudemans, 1929 are both considered to be junior synonyms of D. crassipes Kramer, 1898.


Lectotypes are designated for the following species: D. amaniensis (Vitzthum, 1919), D. braunsi (Vitzthum, 1914), D. brevihirtus LeVeque, 1930, D. crassipes Kramer, 1898, D. inflatus LeVeque, 1930, D. kerrianus LeVeque, 1931, D. maxima (Vitzthum, 1919), D. oudemansi LeVeque, 1930, Dolaea vitzthumi Oudemans, 1926, Dinogamasus philippinensis LeVeque, 1930, D. piperi LeVeque, 1930, Dolaea schoutedeni Oudemans, 1929, Greenia sjoestedti Trägårdh, 1904, Dinogamasus sternisetosa (Vitzthum, 1930).


The genus now includes 36 species (or 37 if D. inflatus LeVeque turns out to be a separate species from Greenia hirtissima Berlese). Based on a phylogenetic analysis, it is discussed whether the genus originated in the far east Asia and that species have invaded Africa on at least two occasions. The phylogenetic analysis is compared to that of the Large Carpenter Bees and, with a few interesting divergences, an overall correspondence between the two systems is found.


ESS 55.
KOMAI, F.. A taxonomic revision of the genus Grapholita and allied genera (Lepidoptera: Tortricidae) in the Palaearctic region1999. 226 pp. SEK 320.

The tribe Grapholitini in the Palaearctic region is revised on the basis of a phylogenetic analysis, which has resulted in recognizing three main genus groups defined here, the Dichrorampha genus group, the Cydia genus group, and the Grapholita genus group. Synapomorphic characters are provided to support the monophyly of the genus groups as well as that of the tribe Grapholitini. The Grapholita genus group has been studied in detail and the phylogenetic relationships among genera have been resolved as far as possible. The Grapholita genus group includes the following eleven genera: Thaumatotibia Zacher, gen. rev., Cryptophlebia Walsingham, Matsumuraeses Issiki, Selania Stephens, Grapholita Treitschke, Pammene Hübner, Dierlia Diakonoff, Pseudopammene Komai, Parapammene Obraztsov, Strophedra Herrich-Schäffer, and Andrioplecta Obraztsov.

Ecological diversification such as geographical distribution, host plant preferences, larval habits, and sex pheromones are summarized. Each genus is redescribed including synonymy, diagnosis, hosts, distribution, and a list of included species. Five new generic synonyms are proposed: Metriophlebia Diakonoff removed from a synonym of Cryptophlebia as a synonym of Thaumatotibia Zacher; Pammenodes Danilevsky & Kuznetzov, Mimarsinania Koçak, and Diplosemaphora Diakonoff as synonyms of Parapammene; Strophedromorpha Diakonoff as a synonym of Strophedra Herrich-Schäffer. Seven new species are described: Grapholita yasudai Komai, Grapholita okui Komai, Grapholita latericia Komai, Pammene fulminea Komai, Pammene piceae Komai, Parapammene reversa Komai, and Strophedra magna Komai.

Lectotypes are designated for Laspeyresia (Grapholitha) cerasana Kozhanchikov and Pammene tsugae Issiki. One synonym, Cryptophlebia lasiandra (Meyrick), is reinstated as a valid species. The following new combinations are given: Thaumatotibia agriochlora (Meyrick), T. apicinudana (Mabille), T. batrachopa (Meyrick), T. chaomorpha (Meyrick), T. citrogramma (Clarke), T. dolichogonia (Diakonoff), T. ecnomia (Diakonoff), T. encarpa (Meyrick), T. etiennei (Diakonoff), T. eutacta (Diakonoff), T. fulturana (Kuznetzov), T. hemitoma (Diakonoff), T. leucotreta (Meyrick), T. macrogona (Diakonoff), T. macrops (Diakonoff), T. nannophthalma (Diakonoff), T. nythobia (Clarke), Selania aeologramma (Meyrick), S. bengalica (Obraztsov), S. diplosperma (Diakonoff), S. exornata (Diakonoff), S. minuta (Obraztsov), Grapholita geministriata (Walsingham), Acanthoclita phaulomorpha (Meyrick), Pammene cyanatra (Diakonoff), P. leucitis (Meyrick), Parapammene amphibola (Diakonoff), P. ellipticopa (Meyrick), P. glaucana (Kennel), P. harmologa (Obraztsov), P. hexaphora (Meyrick), P. petulantana (Kennel), Strophedra dicastica (Meyrick), S. homotorna (Meyrick), S. mica (Diakonoff), S. graphologa (Diakonoff), and S. pericapna (Diakonoff).


ESS 56.
Adam, I. J. & Saether, O. A. Revision of the genus Nilothauma Kieffer, 1921 (Diptera Chironomidae). 1999. 107 pp. SEK 200.

A revised generic diagnosis of the genus Nilothauma Kieffer, 1921a, is given. Toyayusurika Sasa, Suzuki et Sakai, 1998, is a junior synonym of Nilothauma, and T. simantofea Sasa, Suzuki et Sakai, a junior synonym of N. hibaratertia Sasa. N. jintuprimum (Sasa) is shown to be a junior synonym of N. japonicum Niitsuma.


Sixteen new species are described; N. sasai sp. n. from Japan in both sexes and all stages, N. verrucum sp. n. from Canada as 7 and 6 imagines and tentatively associated larva, N. adunatum sp. n. from Australia as 7 imago based on a mature pupa, N. fuscina sp. n., N. duminola sp. n., N. insolitum sp. n., N. flabellatum sp. n., and N. ankasense sp. n. from Ghana, N. burmeisteri sp. n. from Ghana and D. R. Congo, N. mergae sp. n. from Thailand, N. infissum sp. n. from Australia, N. acre sp. n. from China, N. latocaudatum sp. n. from Zimbabwe, N. anderseni sp. n. from Tanzania, N. harrisoni sp. n. from South Africa, and N. kakumense sp. n. from Ghana as 7 imagines. N. japonicum Niitsuma and N. brayi (Goetghebuer) are described or redescribed in both sexes and all stages; N. bicorne (Townes) as 7 imagines, pupae and larvae; N. pictipenne Kieffer as 7 and 6 imagines; N. babiyi (Rempel) and N. mirabile (Townes) as 7 imagines and tentatively associated larvae; N. hibaratertium Sasa, N. nojirimaculatum Sasa and N. hibaraquartum Sasa as 7 imagines. An unassociated pupa from Australia is described. — Keys to 7 imagines, known 6 imagines, known pupae and larvae of Nilothauma are given and the phylogenetic relationships of the Nilothauma group of genera (Nilothauma, Neelamia Soponis, 1986, and Paranilothauma Soponis, 1986) as well as species of Nilothauma discussed.


The genus Paratendipes Kieffer apparently forms the sister group of a group consisting of the genera Paranilothauma, Neelamia and Nilothauma. Within Nilothauma, 4 species groups are recognised with the duminola-group forming the sister group of the remaining groups. The babiyi-group apparently forms the sister group of the brayi and the pictipenne-groups combined. — Paratendipes is cosmopolitan. The Neotropical group of genera (Neelamia and Paranilothauma), probably together with some basal species of Nilothauma, appear to have split off very early with the fragmentation of Pangaea. The species of Nilothauma treated here are of Megagaeic lineages and are confined to the Holarctic, Sino-Indian, Afrotropical and Australian zoogeographical regions for freshwater fauna.


The duminola-group is made up of 2 Afrotropical species. The sister species N. duminola and N. fuscina are exclusively Afrotropical and from the same locality in Ghana. The babiyi-group is made up of 2 Nearctic species which, however, probably originated in the original Gondwanian part of North America. The brayi-group may be divisible in 2 groups. One group includes the Afrotropical N. burmeisteri and N. ankasense, the East Asian N. japonicum and N. mergae and the Nearctic N. bicorne, while in the other group two Australian species (N. adunatum and N. infissum) form the sister group of one European (N. brayi), 2 East Asian (N. sasai and N. acre), one East Asian and probably European (N. hibaratertium), and one Nearctic species (N. mirabile). The pictipenne-group is made up of 2 species (N. nojirimaculatum Sasa and N. hibaraquartum Sasa) from the Sino-Indian region and 7 species (N. insolitum, N. harrisoni, N. anderseni, N. latocaudatum, N. pictipenne, N. flabellatum and N. kakumense) from the Afrotropical Region.


— The phyletic interrelationship of the Nilothauma group of genera combined with the corresponding area cladogram, BPA analysis and estimations of ancestral areas suggests the early splitting of their common ancestor into a Neotropical group (Neelamia, Paranilothauma and perhaps the babiyi-group) and a Megagaeic lineage. The Megagaeic lineage group made up of most Nilothauma species shows a warm/eurythermic vicariant Gondwanan pattern with multiple sister group relationships between the East Asia subregion and the Afrotropical region.


The linkage with the Australian region could be due to the same pattern or, more likely, to recent dispersals from Asia, i.e. post-Miocene contacts. The East Asia Ð North America linkage likely is of post-Miocene origin. The European species, N. brayi and N. ?hibaratertium, most likely is a result of dispersal from East Asia. — The distributions and ecologies of the different species of the genus are outlined.


ESS 57.
Brake, I.: Phylogenetic systematics of the Milichiidae (Diptera, Schizophora). 2000. 120 pp.
SEK 200.

A phylogenetic tree based on adult characters is proposed for the Milichiidae. The computer-based cladistic analysis employed 23 taxa and 53 characters. The monophyly of the Milichiidae and of the subfamily Milichiinae is confirmed. However, the subfamily Madizinae in its former sense is para-phyletic and has therefore been restricted to the genera
Desmometopa, Leptometopa, Litometopa, and Madiza.

These genera are more closely related to the Milichiinae than to the genera Aldrichiomyza, Costalima, genus nov. 1, Microsimus, Neophyllomyza, Paramyia, Phyllomyza, Stomosis, and Xenophyllomyza. For these latter genera, the new subfamily Phyllomyzinae is established. The sister-group of the Milichiidae is the Chloropidae. It is suggested that the Acartophthalmidae is the sister-group of the Milichiidae + Chloropidae, based on the elongation and coiling up of the spermathecal ducts.

The Acartophthalmidae has been used to represent the outgroup for the computer-based cladistic analysis. The morphology and character transformations within the Milichiidae are discussed and the stem-species pattern of the Milichiidae is compared with the stem-species pattern of the Chloropidae, Acartophthalmidae, Carnidae, and the Chloropidae family-group.


The main emphasis is laid on the morphology of the labella, ovipositor, female reproductive system, and male genitalia. A catalogue of world Milichiidae and a key to genera are given.


ESS 58.
Adamski, D. & Brown, J. W.: Systematic revision of the Ecdytolopha group of genera (Lepidoptera: Tortricidae: Grapholitini) in the New World. 2001. 86 pp. SEK 240.

The New World genera Ecdytolopha Zeller, Gymnandrosoma Dyar, revised status, and Pseudogalleria Ragonot are revised. As currently defined, Ecdytolopha includes 10 species: E. insiticiana Zeller, 1875 (type species), ranging from southern Canada to Florida, and from the East Coast to California; E. mana (Kearfott, 1907) from the southern United States (Arizona to Alabama, north to Maryland); E. exploramae, new species, from Peru; E. beckeri, new species, from Tamaulipas, Mexico; E. occidentana, new species, from Arizona, USA; E. coloradana, new species, from Colorado and New Mexico, USA; E. sinaloana, new species, from Sinaloa, Mexico; E. leonana, new species, from Nuevo Leon, Mexico; E. holodesma (Walsingham, 1914), new combination, from Morelos, Hidalgo, and Distrito Federal, Mexico; and E. ricana, new species, from Costa Rica. Gymnandrosoma includes 7 species: G. punctidiscanum Dyar, 1904 (type species), revised status, from the eastern and southeastern USA; G. cryptotortanum, new species, from Veracruz, Mexico; G. linaresensis, new species, from Nuevo Leon, Mexico; G. trachycerus Forbes, 1931, revised status, from the Caribbean; G. leucothorax, new species, from the Caribbean; G. aurantianum Costa Lima, 1927, revised status, from South America, Central America, and the Caribbean; and G. desotanum Heinrich, 1926, revised status, from Florida, USA. Pseudogalleria is monotypic, with the single species P. inimicella (Zeller, 1872), from eastern USA.


Members of these three genera include pests of Citrus, Robinia, Macadamia, Pithecellobium, Theobroma, and Psidium. We present a hypothesis of the phylogenetic relationships among these three New World genera and the Old World genera Cryptophlebia Walsingham and Thaumatotibia Zacher.


ESS 59.
Rognes, K.: Blowflies (Diptera, Calliphoridae) of Israel and adjacent areas, including a new species from Tunisia. 2002. 148 pp. SEK 200.

Calliphoridae of Israel and adjacent areas (Gaza Strip, West Bank, Golan heights, and the Sinai Peninsula) are revised. 43 species in 12 genera occur in the area.


A new species from Tunisia is included, and thus 44 species are dealt with. Ten species are described as new to science, one in the subfamily Calliphorinae, Bellardia nova n. sp., and nine in the subfamily Rhiniinae, Cosmina fishelsohni n.sp., Rhyncomya aravaensis n.sp., R. cyanicolor n.sp., R. negevi n.sp., R. paralutea n.sp., R. sinaiensis n.sp., R. subspeciosa n.sp., R. tunisia n.sp., and R. yahavensis n.sp.


Keys to subfamilies, genera and species are provided. Male and female genitalia are described and illustrated for all new species, for all other Rhiniinae recorded from the area covered (except those figured satisfactorily elsewhere), for Chrysomya albiceps and C. marginalis, for Pollenia dasypoda Portschinsky, and for Bellardia tatrica (Enderlein).


The nominal genus Metallea Wulp is sunk as a junior synonym of Rhyncomya Robineau-Desvoidy, n. syn. The nominal species R. koschewnikovi Rohdendorf, 1930 is sunk as a synonym of R. callopis (Loew) n. syn. A neotype is designated for the nominal species Idia speciosa Loew, 1844 (now in Rhyncomya). The holotype of Idia cyanescens Loew, 1844, is re-discovered and labelled. A lectotype is designated for Onesia fulviceps Egger, 1855.


Three rhiniine species, i.e. R. callopis (Loew), R. sinaiensis n.sp. and R. yahavensis n.sp., are established to have a macrolarviparous reproduction, the first time that such a life history has been reported in Rhiniinae. In two of these species, a large first instar larva was recovered from the female abdomen which it filled for the whole length. The first instar larva of R. sinaiensis n.sp. is described and figured.


These three species plus R. negevi n. sp. form a callopis-species group, all having strong spines along the lower edge of the surstylus. Two rhiniine species, i.e. R. aravaensis n.sp. and R. yahavensis n.sp., are bred from moth larvae. Cosmina arabica Robineau-Desvoidy and Rhyncomya impavida (Rossi) are removed from the list of Calliphoridae from Israel and adjacent areas.


ESS 60.
Johanson, K. A. Systematic revision of the American Helicopsyche of the subgenus Feropsyche (Trichoptera, Helicopsychidae). 2002. 147 pp. SEK 200.

Sixty-four of the seventy-two American Helicopsyche von Siebold, 1856, species within Feropsyche Johanson, 1998, are re-described based on type material and non-type material when types are unavailable.


Reproductions of the original descriptions of the remaining eight species are given in order to present a comprehensive overview of the subgenus in America. Helicopsyche woytkowskii Ross, 1956, earlier synonymized with Helicopsyche peruana Banks, 1920 is considered a valid species. Helicopsyche falcigona Botosaneanu & Flint, 1991a, earlier synonymized with H. ochthephlebia Flint, 1968 is considered a valid species.


Helicopsyche molesta Botosaneanu in Botosaneanu & Hyslop, 1998 is considered a good species. Helicopsyche margaritensis Botosaneanu, 1959, is considered a junior synonym under Helicopsyche vergelana Ross, 1956. Valvata agglutinans Lechmere Guppy, 1864 is considered a synonym under Helicopsyche maculisternum Botosaneanu & Alkins-Koo, 1993.


A key to the males is presented, and maps showing the distribution of all examined species based on the examined material is given.


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